PSS is awesome software to play with brushes setting, but I think it missed something that I think important, it's Initial Direction
. It's mean that the transformation of brush will only changed at initial brush movement .
for example on angle , when we set the angle setting to Initial direction, the brush will only change the angle as long the pen doesn't pulled up ,
maybe it's pretty similar with tilt but keep in mind that not all pen tablet support tilt pen